Because of the detail in the skin texture, I ended up doing a lot more semi-drybrushing than I've done in a very long time.
A closer shot lets you see just how much texture is on the skin.
Finally, here are some shots of the skin after the last highlight. I've tried really hard to preserve blackline areas, and have pulled away from my typical severe highlights by narrowing my dark-to-light range.
A final shot of the wings.
Next, I started on the bone. I've painted so much bone on this blog, that I didn't take any pictures in progress and just jumped straight to the end result: